Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Glandirana emeljanovi (Nikolskii, 1913)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Glandirana > Species: Glandirana emeljanovi

Rana emeljanovi Nikolskii, 1913, Annu. Mus. Zool. Acad. Imp. Sci. St. Petersbourg, 18: 148. Holotype: SZKU, according to Nikolskii, 1918, Fauna Rossii, Zemnovodnye: 85; Stejneger, 1925, Proc. U.S. Natl. Mus., 66: 26. Type locality: Il'yampo [= Imienpo Station], Chinese Eastern Railway, Manchuria [= Yimianpo, Shangzhi County, Heilongjiang Province, China].

Rana rugosa emeljanovi — Terentjev and Chernov, 1940, Kratikii Opredelitel' Presmyka iushchisksia i Zemnovodnykh CCCP: 47.

Rana rugosa emeljanowi — Bannikov, Darevsky, Eshchenko, Rustamov, and Shcherbak, 1977, Operd. Zemn. I Presm. Fauny SSSR: 57. Incorrect subsequent spelling.

Rugosa emeljanowi — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 146; Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 256.

Rana (Rugosa) emeljanovi — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 332.

Rugosa emeljanovi — Fei, 1999, Atlas Amph. China: 184. Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 113; Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 37. See comment under Ranidae record.

Glandirana emeljanovi — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 368.

Common Names

Imienpo Station Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 107).

Northeast China Rough-skinned Frog (Fei, 1999, Atlas Amph. China: 184).

Wrinkled Frog (Borzée, 2024, Continental NE Asian Amph.: 104).

Distribution

Korean Peninsula and northeastern China from southeastern Heilongjiang to southeastern Liaoning; possibly into adjacent Far East Russia.

Geographic Occurrence

Natural Resident: China, People's Republic of, Korea, Democratic People's Republic (North), Korea, Republic of (South)

Likely/Controversially Present: Russia

Comment

Removed from the synonymy of Rana rugosa by Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 332, and Fei, 1999, Atlas Amph. China: 184, where it had been placed by Liu and Hu, 1961, Tailless Amph. China: 141, and Bannikov, Darevsky, Eshchenko, Rustamov, and Shcherbak, 1977, Operd. Zemn. I Presm. Fauny SSSR: 57. The recognition of this taxon is supported by the allozymic results of Yang, Min, Kim, Suh, and Kang, 2000, Korean J. Biol. Sci., 4: 23–30, who noted that Korean and Japanese populations of "Rana rugosa" are distinct species. Yang, Kim, Min, and Suh, 2001, Monogr. Korean Amph.: 70–71, provided a brief account, figure, and map for South Korea (as Rana rugosa). Kuzmin and Maslova, 2003, Adv. Amph. Res. Former Soviet Union, 8: 343–344, discussed this taxon (as Rana rugosa emeljanovi) and noted that no specimen exists for Russia, although there is a chance that it occurs in the Korean border area. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1090–1095, provided an account (as Rugosa emjanovi), figures, and map for China. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 294–295, provided a brief account(as Rugosa emeljanovi) including photographs of specimens and habitat. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 332–333, provided an account (as Rugosa emeljanovi), photographs, and a range map for China that suggests its absence in Russia. Li, Lian, and Lu, 2010, Res. Tadpoles Liaoning: 51–54, described and pictured the larva. Dot map and discussion of status for North Korea, as Rana rugosa, provided by Song, 2016, J. Natl. Park Res., Rep. Korea, 7: 211–217. Liu, Tao, Wang, Zhao, and Liu, 2017, MtDNA, Part B, 2: 383–384, reported on the complete mtDNA genome. Eo, Lee, Park, Jung, Hong, and Lee, 2019, MtDNA, Part B, 4: 961–962, noted on the basis of the complete mtDNA genome that the Korean and Chinese populations are conspecific and distinct from Glandirana rugosa. Borzée, Litvinchuk, Ri, Andersen, Nam, Jon, Man, Choe, Kwon, Othman, Messenger, Bae, Shin, Kim, Maslova, Luedtke, Hobin, Moores, Seliger, Glenk, and Jang, 2021, Animals, 11 (2057): 1–37, provided locality records, a distribution map as well as modeled distribution, life history comments, and conservation status for P.D.R. Korea. Andersen, Chuang, Choe, Kim, Kwon, Jang, and Borzée, 2022, Zool. Stud., Taipei, 61(25): 1–10, reported on the elevational range (0–1198 m) in Rep. Korea. Borzée, 2024, Continental NE Asian Amph.: 104–108, provided a detailed account (distribution including a polygon map, habitat, ecology, conservation, photos of larvae and adults, identification tools) for northeastern Asia.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.