Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hylarana macrodactyla Günther, 1858

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Hylarana > Species: Hylarana macrodactyla

Hylarana macrodactyla Günther, 1858, Arch. Naturgesch., 24: 323. Syntypes: BMNH (6 specimens), by original designation. These may include 1851.12.27.355, 1856.11.17.43–44, and 1933.3.7.39–40, according to museum records but this requires confirmation. Type locality: "China", later given as "Hong Kong" and "China" by Günther, 1859 "1858", Cat. Batr. Sal. Coll. Brit. Mus.: 72, and Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 55; restricted to "Hong Kong", China, by Gee and Boring, 1929, Peking Nat. Hist. Bull., 4: 31, and Taylor, 1962, Univ. Kansas Sci. Bull., 43: 421–423.

Rana trivittata Hallowell, 1861 "1860", Proc. Acad. Nat. Sci. Philadelphia, 12: 504. Holotype: deposition not stated; presumably originally in ANSP or USNM. Type locality: "Hong Kong", China. Synonymy by Boettger, 1888, Ber. Offenbach. Ver. Naturkd., 1888: 96; Boulenger, 1912, in Robinson (ed.), Vert. Fauna of the Malay Peninsula: 238; and (provisionally) by Dubois and Ohler, 2000, Alytes, 18: 32. Formerly considered a possible synonym of Rana plancyi by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 37.

Hylorana macrodactyla — Günther, 1864, Rept. Brit. India: 424; Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 145; Bourret, 1942, Batr. Indochine: 98; Song, Jang, Zou, and Shi, 2002, Herpetol. Sinica, 9: 71; Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 237; by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 368; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 3, by implication.

Hylorana subcoerulea Cope, 1868, Proc. Acad. Nat. Sci. Philadelphia, 20: 139. Syntypes: MCZ 624–626, according to the description; including MCZ 1591, according to Barbour and Loveridge, 1929, Bull. Mus. Comp. Zool., 69: 282. Type locality: "Rangoon, Burmah". Synonymy (with Rana erythraea) by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 65. Subsequently considered a possible synonym of Rana macrodactyla by Boulenger, 1890, Fauna Brit. India, Rept. Batr.: 455; and Boulenger, 1920, Rec. Indian Mus., 20: 155.

Rana macrodactyla — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 54.

Rana (Hylorana) macrodactyla —Boulenger In Mason, 1882, Burma, Ed. 3: 499; Boulenger, 1920, Rec. Indian Mus., 20: 127-130.

Rana (Limnodytes) macrodactyla — Bourret, 1927, Fauna Indochine, Vert., 3: 264.

Rana (Hylarana) macrodactyla — Bourret, 1941, Annexe Bull. Gen. Instr. Publique, Hanoi, 1941: 26; Dubois, 1987 "1986", Alytes, 5: 42, by implication.

Hylarana (Tenuirana) macrodactyla — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 143; Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 122.

Rana (Hylarana) macrodactyla — Ohler and Mallick, 2003 "2002", Hamadryad, 27: 65.

Hylarana (Hylarana) macrodactyla — Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1186.

English Names

Guangdong Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 108).

Long-toed Slender Frog (Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 125; Fei, 1999, Atlas Amph. China: 180).

Marbled Slender Frog (Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 125).

Long-toed Frog (Karsen, Lau, and Bogadek, 1986, Hong Kong Amph. Rept.: 26 Neang and Holden, 2008, Field Guide Amph. Cambodia: 80).

Three-striped Grass Frog (Karsen, Lau, and Bogadek, 1986, Hong Kong Amph. Rept.: 26).

Brown Grasspound Frog (Nutphund, 2001, Amph. Thailand: 114).

Stripe-backed Frog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 138; Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 340).

Long-toed Grass Frog (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 47).

Long-toed Copper-backed Frog (Zug, 2022, Smithson. Contrib. Zool., 653: 37).

Distribution

Southern and northeastern Guangdong, central Guangxi, and Hainan and Hong Kong (China) and Indochina throughout Vietnam, Cambodia, Thailand to and throughout Myanmar and south to northern West Malaysia.

Geographic Occurrence

Natural resident: Cambodia, China, People's Republic of, Hong Kong, Laos, Macao, Malaysia, Malaysia, West (Peninsular), Myanmar, Thailand, Vietnam

Comment

Discussed by Boulenger, 1920, Rec. Indian Mus., 20: 155; Pope, 1931, Bull. Am. Mus. Nat. Hist., 61: 540–542; Bourret, 1942, Batr. Indochine: 338–341; Taylor, 1962, Univ. Kansas Sci. Bull., 43: 421–423; Berry, 1975, Amph. Fauna Peninsular Malaysia: 76–77; Liu and Hu, 1961, Tailless Amph. China: 197–198; and Fei, 1999, Atlas Amph. China: 180–181. Karsen, Lau, and Bogadek, 1986, Hong Kong Amph. Rept.: 26, provided a brief account. See brief account and photo by Matsui, 1979, Contrib. Biol. Lab. Kyoto Univ., 25: 298–300, and Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 112–113. Zhang and Wen, 2000, Amph. Guangxi: 91, provided an account for Guangxi. Reported for southwestern Cambodia by Ohler, Swan, and Daltry, 2002, Raffles Bull. Zool., 50: 465–481. Nutphund, 2001, Amph. Thailand: 114, provided a brief characterization and photograph. Stuart, 1999, in Duckworth et al. (eds.), Wildlife in Lao PDR: 48, commented briefly on the range in Laos. Chan-ard, 2003, Photograph. Guide Amph. Thailand: 138–139, provided a very brief account, map for Thailand, and photograph. Teynié, David, Ohler, and Luanglath, 2004, Hamadryad, 29: 37, provided a record and comments for Laos. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 29, provided specific localities for Vietnam. Stuart, Sok, and Neang, 2006, Raffles Bull. Zool., 54: 140–141, provided a record for Cambodia. Neang and Holden, 2008, Field Guide Amph. Cambodia: 80, provided a photograph, brief account of identification, ecology, and range in Cambodia. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1186–1191, provided an account, figures, and map for China. Chan, Belabut, and Ahmad, 2010, Russ. J. Herpetol., 17: 202–206, considered the species to be part of the West Malaysia fauna but did not mention vouchers. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 304, provided a brief account including photographs. Chan-ard, Cota, and Makchai, 2011, Amph. E. Region Thailand: 102–103, provided a photograph and brief account for the Eastern Region of Thailand. Shi, 2011, Amph. Rept. Fauna Hainan: 115–116, provided an account for Hainan as Rana macrodactyla. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 341, provided an account, photographs, and a range map for China. Wangyal, 2013, J. Threatened Taxa, 5: 4774–4783, reported Hylarana taipehensis from Bhutan (Samdrupjongkhar District), but given the revision of Hasan et al. (2019), the likelihood is that this is referrable to Hylarana tytleri. Minimally the identification needs confirmation. See account, photograph, and map for Vietnam in Vassilieva, Galoyan, Poyarkov, and Geissler, 2016, Photograph. Field Guide Amph. Rept. Lowland S. Vietnam: 103–104. Li, 2011, Amph. Rept. Guangdong: 49, provided a brief account for Guangdong, China, and photograph. Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 340–341, provided a brief account (description, photographs, habitat, and range) for Thailand (in Thai). Makchai, Chuaynkern, Safoowong, Chuachat, and Cota, 2020, Amph. N. Thailand: 134, provided a brief account, photographs, and a range map for Thailand that shows it throughout. Zug, 2022, Smithson. Contrib. Zool., 653: 37–38, briefly discussed identification, habitat, and range in Myanmar. Laurence, Nicolas, Ayoro, and Ohler, 2023, Zootaxa, 5258: 99–112, provided a dot map and discussed distribution modelling and the effect of misidentifications on those models.

External links:

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For additional sources of information from other sites search Google
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.