Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Polypedates mutus (Smith, 1940)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Polypedates > Species: Polypedates mutus

Rhacophorus mutus Smith, 1940, Rec. Indian Mus., 42: 473. Syntypes: BMNH 1940.6.1.3–4, by original designation, now renumbered 1947.2.8.21–22 according to museum records. Type locality: "N'Chang Yang", northern Myanmar.

Polypedates mutus — Inger, 1985, in Frost (ed.), Amph. Species World: 541; Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 184; Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 326.

Rhacophorus mutus — Tian, Jiang, Wu, Hu, Zhao, and Huang, 1986, Handb. Chinese Amph. Rept.: 64.

Rhacophorus (Rhacophorus) mutus — Dubois, 1987 "1986", Alytes, 5: 77.

English Names

Burmese Whipping Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 113).

Vocal Sacless Treefrog (Fei, 1999, Atlas Amph. China: 274).

Northern Treefrog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 152).

Smith's Striped Treefrog (Zug and Mulcahy, 2020 "2019", Amph. Rept. S. Tanintharyi: 63).

Upland Treefrog (Zug, 2022, Smithson. Contrib. Zool., 653: 44).

Distribution

Northern Myanmar (Bago, Kachin, and Shan); isolated populations in southern Yunnan, Guizhou, Guangxi, Guangdong, and Hainan I., China; throughout Vietnam from the border of China to the northern reaches of the Annam mountains in Nghe An Province; northern Laos; uplands of northern and northeastern Thailand (see comment); reported from southern Tanintharyi, peninsular Myanmar (see comment); expected from Guangxi, China.

Comment

In the Rhacophorus (Rhacophorus) leucomystax group of Dubois, 1987 "1986", Alytes, 5: 77; see comment under Polypedates. Removed from the synonymy of Polypedates leucomystax by Liu and Hu, 1961, Tailless Amph. China: 248–250. Yang, 1991, Amph. Fauna of Yunnan: 208–210; and Fei, 1999, Atlas Amph. China: 274–275, provided accounts. Orlov, Lathrop, Murphy, and Ho, 2001, Russ. J. Herpetol., 8: 36–38, provided an account for the Vietnam population. Zhang and Wen, 2000, Amph. Guangxi: 129, provided an account for Guangxi. Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 96, reported the species to be in Laos. Li and Hu, 1996, Zool. Res., Kunming, 17: 483–488, reported on karyology. See also brief account by Zhao and Yang, 1997, Amph. Rept. Hengduan Mountains Region: 125–126. Chan-ard, 2003, Photograph. Guide Amph. Thailand: 152–153, provided a very brief account, map for Thailand, and photograph. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 36, provided specific localities for Vietnam and (p. 150) a photograph. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 111, provided a brief account for Yunnan, China. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 760–764, provided an account and a spot map for China. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 448, provided a brief account for China including photographs of specimens. Shi, 2011, Amph. Rept. Fauna Hainan: 66–69, provided an account for Hainan. Kuraishi, Matsui, Hamidy, Belabut, Ahmad, Panha, Sudin, Yong, Jiang, Ota, Thong, and Nishikawa, 2013, Zool. Scripta, 42: 54–70, reported on the phylogenetics of the Polypedates leucomystax group and referred to two species as Polypedates cf mutus 1 and Polypedates cf mutus 2 pending investigation of the population at the type locality. The current stated range is, of course, highly provisional as a result of this finding. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 530, provided an account, photographs, and a range map for China. Hecht, Pham, Nguyen, Nguyen, Bonkowski, and Ziegler, 2013, Biodiversity J., 4: 507–552, reported a record from Tay Yen Tu Nature Reserve, Bac Giang Province, northeastern Vietnam and discussed the range. Pan, Dang, Wang, Zheng, Rao, and Li, 2013, Asian Herpetol. Res., 4: 124–133, provided a phylogeny reconstruction of the Polypedates leucomystax complex that suggests that this nominal species extends to northern Myanmar. Ziegler, Rauhaus, Tran, Pham, van Schingen, Dang, Le, and Nguyen, 2015, Sauria, Berlin, 37: 11–44, provided a locality in Vin Phuc Province, Vietnam. Pham, An, Herbst, Bonkowski, Ziegler, and Nguyen, 2017, Bonn Zool. Bull., 66: 37–53, provided records for Cao Bang Province, Vietnam, along with observations on morphology and natural history. Li, 2011, Amph. Rept. Guangdong: 70, provided a brief account for Guangdong, China, and photograph. Zhang, 2017, Amph. Rept. Fanjing Mts.: 165–167, provided taxonomic and natural history information for the Fanjing Mountains population in northeastern Guizhou, China. Poyarkov, Nguyen, Duong, Gorin, and Yang, 2018, PeerJ, 6 (e5771): 17, provided a record for Cat Ba National Park, Hai Phong Province, Vietnam. Mulcahy, Lee, Miller, Chand, Thura, and Zug, 2018, ZooKeys, 757: 95, provided a genetically confirmed record from Taninthary Division, southern Myanmar. Liu, Wang, Zhai, and Wang, 2019 "2018", Asian Herpetol. Res., 9: 240–249, reported on call characteristics. Reported from Lai Chau Province, Vietnam, by Pham, Hoang, Pham, and Nguyen, 2019, VNU J. Sci. Nat. Sci. Tech., 35: 73–79. Zug and Mulcahy, 2020 "2019", Amph. Rept. S. Tanintharyi: 63–64, provided a brief account for South Tanintharyi, peninsular Myanmar, but noted that it is unlikely that this population is conspecific with the northern Myanmar populations. See also Sutthiwisesa, Taksintum, Arunyawat, Sangthong, and Jantrarotai, 2020, Agr. Nat. Res., Thailand, 54: 1–8, reported at least two species lineages under this name in Thailand. Pham, Do, Ngo, Tran, Ziegler, and Nguyen, 2020, Check List, 16: 1025–1041, provided a record from Hai Ha District, Quang Ninh Province, Vietnam, on the Guangxi, China, border. Makchai, Chuaynkern, Safoowong, Chuachat, and Cota, 2020, Amph. N. Thailand: 165, provided photographs, a brief account for Thailand, and a range map. Zug, 2022, Smithson. Contrib. Zool., 653: 44, briefly discussed habitat, range, and identification in Myanmar.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist; for a quick link to their maps see iNaturalist KML
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.