Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Odorrana tiannanensis (Yang and Li, 1980)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Odorrana > Species: Odorrana tiannanensis

Rana tiannanensis Yang and Li, 1980, Zool. Res., Kunming, 1: 261. Holotype: KIZ 77I0185, by original designation. Type locality: "Hekou, Dawei Hill, Yunnan; altitude 1200 m", China.

Odorrana tiannanensis — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 149; Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 261; Song, Jang, Zou, and Shi, 2002, Herpetol. Sinica, 9: 70.

Rana (Odorrana) tiannanensis — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 329.

Rana (Odorrana) megatympanum Bain, Lathrop, Murphy, Orlov, and Ho, 2003, Am. Mus. Novit., 3417: 50. Holotype: ROM 39684, by original designation. Type locality: "Khe Moi River, approximately 24 km west of Con Cuong village (by road), Con Cuong District, Nghe An Province, Vietnam (18° 56′ 30″ N, 104° 48′ 35″ E)". Synonymy by Ohler, 2007, Alytes, 25: 59.

Rana tabaca Bain and Nguyen, 2004, Am. Mus. Novit., 3453: 17. Holotype: AMNH 163923, by original designation. Type locality: "from a stream near the Hmong Village of Khau Ria, in Du Gia commune, Yen Minh District, below Mount Muong Cha, Ha Giang Province, Vietnam (22° 54′ 27″ N, 105° 13′ 59″ E, ca. 800 m elevation)". Synonymy with Rana megatympanum by Stuart and Bain, 2005, Herpetologica, 61: 478-492; with Rana tiannanensis by Ohler, 2007, Alytes, 25: 59.

Odorrana (Odorrana) tiannanensis — Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 130.

Odorrana megatympanum — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 239; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1-13; by implication; Stuart, 2008, Mol. Phylogenet. Evol., 46: 54.

Rana heatwolei Stuart and Bain, 2005, Herpetologica, 61: 487. Holotype: FMNH 258134, by original designation. Type locality: "on a tree root projecting from a dirt bank 2 m above a tributary of the Nam Ou River in evergreen forest in Phou Dendin National Biodiversity Conservation Area, Phongsaly District, Phongsaly Province, Laos, 22° 5′ 38″ N, 102° 12′ 50″ E, 600 m elevation". Synonymy by Ohler, 2007, Alytes, 25: 59.

Huia tiannanensis — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 368.

Huia megatympanum — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 368.

Huia tabaca — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 368.

Odorrana tiannanensis — Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1-13, by implication.

Odorrana heatwolei — Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1-13, by implication.

Odorrana heatwolei — Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 49.

English Names

Dawei Frog (Odorrana tiannanensis: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 109).

Tiannan Odorous Frog (Odorrana tiannanensis: Fei, 1999, Atlas Amph. China: 196).

Megatympanum Frog (Rana megatympanum [no longer recognized]: Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 30).

Tobacco Frog (Rana tabaca [no longer recognized]: Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 31).

Large-eared Odorous Frog (Odorrana megatympanum [no longer recognized]: Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 50).

Distribution

Southern Yunnan (Mangla, Pingbian, and Hekou counties) , China; Phou Dendin National Biodviersty Conservation area, Phongsaly Province, Laos; northern, north-central, and northeastern Vietnam (Ha Giang, Tuyen Quang, Nghe An, Ha Tinh, Quang Binh, Quang Nam, and Binh Dinh Provinces; see comment) in and around montane rivers as well as in the adjacent forest, 120-1000 m elevation

Comment

Section Hylarana, subsection Hylarana, Rana (Odorrana) andersonii group of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 329. Related to Rana anlungensis, according to the original publication. Yang, 1991, Amph. Fauna of Yunnan: 153–155, and Fei, 1999, Atlas Amph. China: 200–201, provided accounts. Ye and Fei, 2001, Acta Zool. Sinica, 47: 528–534, placed this in their Odorrana schmackeri group. In the Odorrana (Odorrana) schmackeri group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 126. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 30, provided specific localities of Rana megatmpanum for Vietnam. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 148, provided a photograph of nominal Rana tabaca. See discussion by Ohler, 2007, Alytes, 25: 55–74, who placed Rana megatympanum and Rana heatwolei into synonymy based on morphological characteristics. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 81–82, provided a brief account (as Rana tiannanensis) for Yunnan, China. See statement of geographic range, habitat, and conservation status (as Rana tiannanensis) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 633, who suggested that the Hainan population may represent a distinct species. Mo, Zhou, Xie, and Chen, 2007, Herpetol. Sinica, 11: 15–18, reported the species in Guangxi, China. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1294–1298, provided an account, figures, and map for China, and placed it in their Odorrana (Odorrana) schmackeri group. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 345, provided a brief account including photographs. Shi, 2011, Amph. Rept. Fauna Hainan: 98–100, provided an account for Hainan as Rana tiannanensis. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 392, provided an account, photographs, and a revised range map for China. Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 49–50, disputed the synonymy of Rana megatympanum and Rana heatwolei for undisclosed reasons.Reported from Son La Province, northwestern Vietnam, by Pham, Nguyen, Pham, Sung, Le, Vaxong, and Ziegler, 2022, Herpetol. Notes, 15: 169–178, who also provided descriptive notes on the new specimens. Duong and Ngo, 2022, Herpetol. Notes, 15: 751–762, discussed range in Vietnam and provided records for Binh Dinh Province (which being in southern Vietnam is a big enough extension to require genetic confirmation: DRF).

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