Zhangixalus dennysi (Blanford, 1881)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Zhangixalus > Species: Zhangixalus dennysi

Rhacophorus dennysi Blanford, 1881, Proc. Zool. Soc. London, 1881: 224. Holotype: ZRC according to the original publication; RMNH, according to Stejneger, 1925, Proc. U.S. Natl. Mus., 66: 31, apparently in error. Type locality: "China; but the precise locality is unknown".

Rhacophorus exiguus Boettger, 1894, Ber. Senckenb. Naturforsch. Ges., 1894: 148. Types: Not stated; SMF 6987 is holotype according to Mertens, 1967, Senckenb. Biol., 48(A): 49. Type locality: "Chinhai bei Ningpo", China. Considered a subspecies of Rhacophorus schlegelii by Wolf, 1936, Bull. Raffles Mus., 12: 194, but this is currently untenable. Synonymy by Liu and Hu, 1961, Tailless Amph. China: 258; Zhao and Adler, 1993, Herpetol. China: 155.

Polypedates dennysiStejneger, 1925, Proc. U.S. Natl. Mus., 66: 31; Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 184.

Hyla albotaeniata Vogt, 1927, Zool. Anz., 69: 287. Holotype: ZMB 24118, according to Pope, 1931, Bull. Am. Mus. Nat. Hist., 61: 462. Type locality: "Südchina". Probably Yunnan Province, according to Zhao and Adler, 1993, Herpetol. China: 155. Synonymy by Liu and Hu, 1961, Tailless Amph. China: 258;.

Polypedates feyi Chen, 1929, China J. Sci. Arts, Shanghai, 10: 198. Holotype: Sun Yat-sen Univ. Collection 1205, by original designation. Type locality: "Yaoshan [= Dayao Shan], Kwangsi [= Guangxi Zhuang Autonomous Region]", China. Synonymy by Pope, 1931, Bull. Am. Mus. Nat. Hist., 61: 563, and Liu and Hu, 1961, Tailless Amph. China: 258.

Rhacophorus (Rhacophorus) exiguusAhl, 1931, Das Tierreich, 55: 113.

Rhacophorus (Rhacophorus) dennysiiAhl, 1931, Das Tierreich, 55: 162; Dubois, 1987 "1986", Alytes, 5: 77.

Rhacophorus nigropalmatus exiguusWolf, 1936, Bull. Raffles Mus., 12: 201.

Rhacophorus nigropalmatus dennysiiWolf, 1936, Bull. Raffles Mus., 12: 201.

Rhacophorus dennysi — Inger, 1966, Fieldiana, Zool., 52: 321; Wilkinson, Drewes, and Tatum, 2002, Mol. Phylogenet. Evol., 24: 272.

Polypedates dennysiLiem, 1970, Fieldiana, Zool., 57: 98.

Rhacophorus (Rhacophorus) dennysiiDubois, 1987 "1986", Alytes, 5: 77.

Zhangixalus dennysi — Jiang, Jiang, Ren, Wu, and Li, 2019, Asian Herpetol. Res., 10: 7. 

English Names

Blanford's Whipping Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 113).

Large Treefrog (Fei, 1999, Atlas Amph. China: 276–277).

Denny's Whipping Frog (Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 36).


Southeastern China from southern Henan and southern Jiangsu south to northern Guangdong, and through central Guangxi to Vietnam; Myanmar; northern and northeastern Vietnam east of the Red River (Cao Bang, Bac Thai, Lang Son, Vinh Phu, Hai Hung, Quang Ninh, and Ha Bac provinces, 200–1500 m elevation); Khammouan Province, Laos; Hainan I., China.


Discussed (as Rhacophorus dennysii) by Liu and Hu, 1961, Tailless Amph. China: 258–261. In the Rhacophorus (Rhacophorus) dennysii group of Dubois, 1987 "1986", Alytes, 5: 77 (see comment under Rhacophorus for dissenting opinion regarding the recognition of this group). See account by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 327; and Fei, 1999, Atlas Amph. China: 277–278 (who excluded China adjacent to Myanmar). Ohler, Marquis, Swan, and Grosjean, 2000, Herpetozoa, Wien, 13: 71–87, reported the Lao Cai Province, Vietnam record. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 204–205, provided a brief account and illustration (as Rhacophorus dennysi). Wilkinson, Drewes, and Tatum, 2002, Mol. Phylogenet. Evol., 24: 265–273, provided molecular and morphological evidence for retaining this species in Rhacophorus and not in Polypedates. Huang, 1990, Fauna Zhejiang, Amph. Rept.: 85–86, provided an account for Zhejiang populations. Zhang and Wen, 2000, Amph. Guangxi: 124, provided an account for Guangxi. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 37, provided specific localities for Vietnam and (p. 150) a photograph. Stuart, 2005, Herpetol. Rev., 36: 478, provided a specific locality for Laos. Orlov, Nguyen, and Ho, 2008, Russ. J. Herpetol., 15: 67–84, provided a photograph and a key to differentiate this species from others in Vietnam. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 789–795, provided an account and a spot map for China and assigned this species to their Rhacophorus maximus group. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 454–455, provided a brief account for China including photographs of specimens. Shi, 2011, Amph. Rept. Fauna Hainan: 65–66, provided an account for Hainan. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 537–538, provided an account, photographs, and a range map. Pham, An, Herbst, Bonkowski, Ziegler, and Nguyen, 2017, Bonn Zool. Bull., 66: 37–53, provided records for Cao Bang Province, Vietnam, along with observatons on morphology and natural history. Shen, 2014, Fauna Hunan, Amph.: 316–321, provided an account for Hunan, China. Li, 2011, Amph. Rept. Guangdong: 71, provided a brief account for Guangdong, China, and photograph. Zhang, 2017, Amph. Rept. Fanjing Mts.: 172–175, provided taxonomic and natural history information for the Fanjing Mountains population in northeastern Guizhou, China. Pham, Do, Ngo, Tran, Ziegler, and Nguyen, 2020, Check List, 16: 1025–1041, provided a record from Hai Ha District, Quang Ninh Province, Vietnam, on the Guangxi, China, border.            

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.