Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nymphargus Cisneros-Heredia and McDiarmid, 2007

Class: Amphibia > Order: Anura > Family: Centrolenidae > Subfamily: Centroleninae > Genus: Nymphargus

43 species

Nymphargus Cisneros-Heredia and McDiarmid, 2007, Zootaxa, 1572: 34. Type species: Nymphargus cochranae (Goin, 1961) [= Cochranella cochranae Goin, 1961], by original designation.

English Names

None noted.

Distribution

Western Andean slopes of Colombia and Ecuador and the eastern Andean slopes of Colombia, Ecuador, Peru, and Bolivia

Comment

Velásquez-Álvarez, Rada, Sánchez-Pacheco, and Acosta-Galvis, 2007, S. Am. J. Herpetol., 2: 191-197, summarized the known ranges of the species of Nymphargus. Guayasamin, Castroviejo-Fisher, Trueb, Ayarzagüena, Rada, and Vilà, 2009, Zootaxa, 2100: 1-97, redelimited the taxon and reported on phylogenetics.

Contained taxa (43 sp.):

Nymphargus anomalus (Lynch and Duellman, 1973)

Nymphargus armatus (Lynch and Ruiz-Carranza, 1996)

Nymphargus balionotus (Duellman, 1981)

Nymphargus bejaranoi (Cannatella, 1980)

Nymphargus buenaventura (Cisneros-Heredia and Yánez-Muñoz, 2007)

Nymphargus cariticommatus (Wild, 1994)

Nymphargus caucanus Rada, Ospina-Sarria, and Guayasamin, 2017

Nymphargus chami (Ruiz-Carranza and Lynch, 1995)

Nymphargus chancas (Duellman and Schulte, 1993)

Nymphargus cochranae (Goin, 1961)

Nymphargus colomai Guayasamin and Hutter, 2020

Nymphargus cristinae (Ruiz-Carranza and Lynch, 1995)

Nymphargus garciae (Ruiz-Carranza and Lynch, 1995)

Nymphargus grandisonae (Cochran and Goin, 1970)

Nymphargus griffithsi (Goin, 1961)

Nymphargus humboldti Guayasamin, Cisneros-Heredia, McDiarmid, and Hutter, 2020

Nymphargus ignotus (Lynch, 1990)

Nymphargus lasgralarias Hutter and Guayasamin, 2012

Nymphargus laurae Cisneros-Heredia and McDiarmid, 2007

Nymphargus lindae Guayasamin, 2020

Nymphargus luminosus (Ruiz-Carranza and Lynch, 1995)

Nymphargus luteopunctatus (Ruiz-Carranza and Lynch, 1996)

Nymphargus manduriacu Guayasamin, Cisneros-Heredia, Vieira, Kohn, Gavilanes, Lynch, Hamilton, and Maynard, 2019

Nymphargus mariae (Duellman and Toft, 1979)

Nymphargus megacheirus (Lynch and Duellman, 1973)

Nymphargus megistus (Rivero, 1985)

Nymphargus mixomaculatus (Guayasamin, Lehr, Rodríguez, and Aguilar, 2006)

Nymphargus nephelophila (Ruiz-Carranza and Lynch, 1991)

Nymphargus ocellatus (Boulenger, 1918)

Nymphargus oreonympha (Ruiz-Carranza and Lynch, 1991)

Nymphargus phenax (Cannatella and Duellman, 1982)

Nymphargus pijao Montilla, Arcila-Pérez, Toro-Gómez, Vargas-Salinas, and Rada, 2023

Nymphargus pluvialis (Cannatella and Duellman, 1982)

Nymphargus posadae (Ruiz-Carranza and Lynch, 1995)

Nymphargus prasinus (Duellman, 1981)

Nymphargus rosada (Ruiz-Carranza and Lynch, 1997)

Nymphargus ruizi (Lynch, 1993)

Nymphargus siren (Lynch and Duellman, 1973)

Nymphargus spilotus (Ruiz-Carranza and Lynch, 1997)

Nymphargus sucre Guayasamin, 2013

Nymphargus truebae (Duellman, 1976)

Nymphargus vicenteruedai Velásquez-Álvarez, Rada, Sánchez-Pacheco, and Acosta-Galvis, 2007

Nymphargus wileyi (Guayasamin, Bustamante, Almeida-Reinoso, and Funk, 2006)

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